Oral Presentation Lorne Infection and Immunity 2014

The mechanisms driving assembly of bacterial cell surfaces. (#20)

Trevor Lithgow 1 , Richard Strugnell 2 , Elizabeth Hartland 2 , Susan Buchanan 3
  1. Monash University, Clayton, VIC, Australia
  2. Department of Microbiology & Immunology, University of Melbourne, Parkville
  3. NIDDK, NIH, Bethesda, USA

In bacteria, outer membrane proteins are essential for nutrient import, signaling, motility and survival. The beta-barrel assembly machinery (BAM) complex is responsible for the biogenesis of beta-barrel membrane proteins, which represent the vast majority of bacterial outer membrane proteins. We solved the structure of the component parts of the BAM complex to determine how it functions. We also discovered a new translocation and assembly module of the machinery (the TAM) which is required for the assembly of many virulence factors in the outer membranes of bacterial pathogens; its structure revealed the way in which it functions together with the core BAM complex. But it has been clear that key virulence factors: such as the type III and type II secretion systems and the capsule secretion system have outer membrane protein portals that are not of beta-barrel architecture. New assays to measure the assembly of these secretion systems demonstrate that a distinct and novel machinery is responsible for their assembly and function in pathogenesis.

  1. Noinaj N, Kuszak AJ, Gumbart JC, Lukacik P, Chang H, Easley NC, Lithgow T, Buchanan SK. (2013) Structural insight into the biogenesis of β-barrel membrane proteins. Nature 501:385-90
  2. Selkrig J, Mosbahi K, Webb CT, Belousoff MJ, Perry AJ, Wells TJ, Morris F, Leyton DL, Totsika M, Phan MD, Celik N, Kelly M, Oates C, Hartland EL, Robins-Browne RM, Ramarathinam SH, Purcell AW, Schembri MA, Strugnell RA, Henderson IR, Walker D, Lithgow T. (2012) Discovery of an archetypal protein transport system in bacterial outer membranes. Nat Struct Mol Biol. 19:506-10